CRU research articles, talks and conference poster presentations are listed below. Many of these citations have weblinks, which will bring you to a post-print article, pdf file, or external website where you can access these materials. If you are having trouble finding a document you need, please feel free to contact us.



Ryan, M.G., Storey, A.E., Anderson, R.E. & Walsh, C.J. (in press, July 2019). Physiological indicators of attachment in domestic dogs (Canis familiaris) and their owners in the Strange Situation Test. Frontiers in Behavioral Neuroscience (Special Issue: The Neurobiology of Human-Animal Bonding). doi:10.3389/fnbeh.2019.00162.


Ryan, M.G., Storey, A.E., Anderson, R.E., & Walsh, C.J. (in revision). Personality traits and owner-dog attachment in a Canadian sample. Human-Animal Interaction Bulletin. Special Issue: Basic Social and Personality Psychology on Human-Animal Interactions.


Walsh, C.J. (2018). Lessons from behaviour for brain imaging. Animal Sentience 22(7).


Howse, M.S., Anderson, R.E., & Walsh, C. J. (2018). Social behaviour of domestic dogs (Canis familiaris) in a public off-leash dog park. Behavioural Processes 157:691-701. doi:10.1016/j.beproc.2018.03.016. PDF (pre-print)


Posluns, J.A., Anderson, R.E., & Walsh, C.J. (2017). Comparing two canine personality assessments: Convergence of the MCPQ-R and DPQ and consensus between dog owners and walkers. Applied Animal Behaviour Science 188: 68-76.


Ottenheimer-Carrier, L., Ricketts, C.J., Perry, E.A., Anderson, R.E., & Walsh, C.J. (2017). Owner-reported personality assessments are associated with breed groups but not with oxytocin receptor gene polymorphisms in domestic dogs (Canis familiaris). Journal of Veterinary Behavior 18: 62-68.


Ottenheimer Carrier, L., Cyr, A., Anderson, R. E., & Walsh, C. J. (2013). Exploring the dog park:

Relationships between social behaviours, personality and cortisol in companion dogs. Applied Animal Behaviour Science, 146 (1-4), 96-106. doi:10.1016/j.applanim.2013.04.002  PDF


Schneider, J. N. & Anderson, R. E. (2011). Tonal vocalizations in the red wolf (Canis rufus): Potential

functions of nonlinear sound production. Journal of the Acoustical Society of America, 130(4),

2275-2284. doi:10.1121/1368331


Anderson, R.E., Russell, J., White, J., & Weir, J. (2001). Hyperwolf 2.0



Graduate Student Theses & Talks


Castro, M. (2017). Assesing the social organization of multi-dog households: Dog behaviour, hormones, personality, and demographics. MSc thesis, Cognitive & Behavioural Ecology Program, Memorial University

of Newfoundland.  

               Link to Thesis; Link to Talk


McDowell, K.E. (2017). Human preferences of canine coat colour and length.  MSc thesis, Cognitive &

Behavioural Ecology Program, Memorial University of Newfoundland.  Thesis            


Posluns, J.A. (2017). Behaviour differences in companion dogs in response to unfamiliar conspecific contact

and personality measusrements.  MSc thesis, Cognitive & Behavioural Ecology Program, Memorial

University of Newfoundland.  

               Link to Thesis  Talk


Howse, M. (2016). Exploring the social behaviour of domestic dogs (Canis familiaris) in a public off-leash dog park. MSc thesis, Cognitive & Behavioural Ecology Program, Memorial University of Newfoundland.       

               Link to Thesis


Ryan, M.G. (2015). Interspecific attachment: Social bonds between humans and their 'best friends'.  MSc

thesis, Cognitive & Behavioural Ecology Program, Memorial University of Newfoundland.  

               Link to Thesis


Posluns, J. (2014). Greetings among unfamiliar dogs: Investigating personality, social behaviour and cortisol.

          Link to Talk given at SPARCS 2014.


Conference Proceedings


Walsh, C.J., Castro, M., Kinnunen, M. & Anderson, R.E. (2019), Resolving dominance in dogs: Behavioral measure fail to predict relationships in mulit-dog homes. Poster presentation at the Animal Behaviour Society Conference, Chicago, IL, USA, July 2019. Poster.


Walsh, C.J., Castro, M., Kinnunen, M., & Anderson, R. E. (2018). The "Dual Hormone Hypothesis"

in domestic dogs: Cortisol X Testosterone interactions predict wins in a competitive task. Poster

presentation at the Canadian Society for Brain, Behaviour, and Cogntive Science, St. John's, NL., Canada, July 2018. Poster.


Walsh, C.J., Castro, M., Kinnunen, M., & Anderson, R. E. (2017). The "dual hormone hypothesis"

in domestic dogs: Interactions of testosterone and cortisol predict competitive outcomes. Oral

presentation at the 1st Canine Science Conference, Tempeh, AZ, USA, October 2017. Talk Slides


Ottenheimer Carrier, L., Ricketts, C. J., Perry, E. A., Anderson, R. E. & Walsh, C. J. (2016). Owner

evaluations of dog personality and behaviour are associated with breed group, but not with two oxytocin

receptor gene polymorphisms. Poster presentation at the 5th Canine Science Forum, Padova,

Italy, July 2016. Abstact: Dog Behavior 3, suppl.: s102. Poster.


Ryan, M. G., Storey, A. E., Anderson, R.E. & Walsh, C.J. (2016). Evidence for synchronization of stress

hormones in owners and dogs during the strange situation test. Poster presentation at the 5th Canine Science Forum, Padova, Italy, July 2016. Abstact: Dog Behavior 3, suppl.: s79. Poster.


Walsh, C.J., Anderson, R.E., Ottenheimer Carrier, L., Penney, J., & Croucher, K. (2014). Canine salivary cortisol in frequent dog park visitors is predicted by individual differences in Neuroticism (MCPQ-R). Journal of Veterinary Behavior, 9, e14. doi:10.1016/j.veb.2014.09.048. Oral presentation at the 4th Canine Science Forum, Lincoln, United Kingdom, July 2014.  Talk Slides (pdf).


Posluns, J., Anderson, R.E., & Walsh, C.J. (2014). Extroverts make new friends: Multiple indicators reflect

successful interactions among unfamiliar dogs. Journal of Veterinary Behavior, 9: e10

Poster presentation at the 4th Canine Science Forum, Lincoln, United Kingdom, July 2014.  Poster


Ottenheimer Carrier, L., Anderson R.E., & Walsh, C.J. (2013). Personality, social behaviour, and cortisol in dogs: Relationships in a dog park. Journal of Veterinary Behavior, 8(4): e43. Oral presentation by C.W. to 3rd Canine Science Forum, Barcelona, Spain, July 25-27, 2012.

     Talk Slides (pdf)


Castro, M. & Walsh, C.J. (2013). Do cortisol and testosterone levels covary with social role in domestic dogs? Journal of Veterinary Behavior, 8(4): e29-30. Poster presentation to 3rd Canine Science Forum, Barcelona, Spain, July 25-27, 2012. Poster Teaser.


Dutton, E. E., Anderson, R. E., Walsh, C. J. (2011). “Do I know you?” Does partner familiarity influence social  interactions among dogs in a park setting? Journal of Veterinary Behavior: Clinical Applications and

Research,6, 65. Poster


Walsh, C. (2011). Dog Parks: What do we know about what dogs do?  

Presentation to IFAAB 2011, New Orleans, USA, Febuary 2011. Abstract   Talk Handout


Walsh, C. J., Howse, M., Green, C., Butler, L., & Anderson, R. E. (2011). “Stop that!”: People interrupting dog

behaviors in a dog park setting. Journal of Veterinary Behavior: Clinical Applications and Research, 6, 77.

      Poster


Howse, M., Walsh, C. J., & Anderson, R. E. (2008). Normal social behaviour of domestic dogs (Canis familiaris). Poster presentation at Animal Behaviour Society Conference, Snowbird, Utah, USA, August 2008. Poster


Completed Honours Theses


Note that Memorial University's Center for Newfoundland Studies archives all dissertations and

theses. Below are links to posters (or talks) which summarize the main findings of the research projects. Many of these presentations were also given by the students at a Science Atlantic undergraduate conference.



White, C. (2019). Assessing the relationship between personality and behavioural responses of domestic dogs during novel object tests. BSc Honours Dissertation, Department of Biology, Memorial University. Thesis


Lamarre, J. (2018). Lifestyles of the modern dog (Canis familiaris): Relationships among activity, feeding,                                                                          personality, and separation anxiety in beagles and huskies. BSc Honours Dissertation, Department of Biology, Memorial University. Thesis


O'Reilly, A. (2018).  The modern dog: Canine metabolic, behavioural, and cognitive indicators of wellness (I).

BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis



Collins, R.  (2017). Dog-owner attachment and proximity behaviours in an off-leash dog park. BSc Honours

Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis


Lambert, G. (2017). How social network metrics among transient grups of dogs relate to physiology and

personality. BSc Honours Dissertation, Department of Biology, Memorial University of Newfoundland. Thesis Presentation


Luther, M. (2017). How initial greetings with conspecifics influence the behavioural and physiological responses of domestic dogs in off-leash dog parks. BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis Poster


Morey, K. (2017). "My dog has separation anxiety": Does owner behaviour influence dog behaviour and hormones during the Strange Situation Test? BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis Presentation


Ricketts, C.J. (2015). Investigating the relationship between oxytocin receptor gene variation and pesonality

in dogs. BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland.  Thesis Poster


O’Leary, J. (2013). Making an entrance: Factors influencing chemosensory investigation patterns in a dog park.

BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis Poster


Clarke, C. (2013). The relationship between coat colour and behavioural problems in Beagle dogs. BSc

Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis Poster


Ottenheimer Carrier, L. (2011). Personality, cortisol levels, and conspecific behaviour in domestic dogs. BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis


Cyr, A. (2011). The relationship between social activities of dogs (Canis familiaris) in a dog park and cortisol.

BSc Honours Dissertation, Department of Psychology, Memorial University of Newfoundland. Thesis Poster


Fraser, K. (2010). Dominance in domestic dogs (Canis familiaris): No evidence for social hierarchies in a transient social group. BSc Honours Dissertation, Department of Psychology, Memorial University of

Newfoundland.   Poster


LINKS